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Pteridaceae
Pteridaceae image
Sue Carnahan
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Michael D. Windham in Flora of North America (vol. 2)
Plants perennial [annual], on rock or terrestrial, of small (rarely large) stature . Stems compact to creeping, branched or unbranched, dictyostelic, bearing hairs and/or scales. Leaves monomorphic to dimorphic, circinate or noncircinate in bud . Petioles usually with persistent scales proximally, lacking spines; vascular bundles 1--several, roundish or crescent-shaped in cross section. Blades 1--6-pinnate, without laminar buds . Indument on petioles, rachises, costae, and blades, rarely absent or commonly of hairs, glands, and/or scales, occasionally of white or yellow farina. Veins pinnate or parallel in ultimate segments of blades, simple or forked, free or infrequently anastomosing in complex patterns. Sori borne abaxially on veins, often confluent with age and forming a continuous submarginal band, or sporangia densely covering abaxial surface (acrostichoid); receptacle not or only slightly elevated. Indusia (when present) formed by reflexed, recurved, or revolute leaf margin (false indusium). Sporangia stalk of 2--3 rows of cells; annulus vertical, interrupted by stalk; spores 64 or 32 (rarely 16) per sporangium. Spores all 1 kind, brown, black, or gray (rarely yellow), globose to globose-tetrahedral or trigonal, occasionally with prominent equatorial ridge, trilete, or trigonal, variously ornamented (usually cristate or rugose). Gametophytes green, aboveground, obcordate to reniform, sometimes asymmetric, usually glabrous (glandular-farinose in Notholaena ); archegonia and antheridia borne on abaxial surface, antheridia 3-celled. Considerable disagreement exists concerning the circumscription and proper name of this family. The taxa comprising the Pteridaceae in this treatment were assigned to the Sinopteridaceae and Pteridaceae by D. B. Lellinger (1985) and were included in five families by R. E. G. Pichi-Sermolli (1977). The broad concept followed here is similar (except for the exclusion of Ceratopteris ) to that espoused by R. M. Tryon and A. F. Tryon (1982), who applied the name Pteridaceae to the group. Until very recently, the newer name Adiantaceae was more commonly used. As represented in North America, Pteridaceae comprise three major evolutionary lines (the adiantoids, the pteroids, and the cheilanthoids). Characteristics holding the family together include abaxial (usually submarginal) sori that lack indusia or are protected by a reflexed or revolute leaf margin, spores that are usually globose-tetrahedral and trilete, and chromosome base numbers of 30 or 29 (rarely 27). The xeric-adapted members of the family (particularly the cheilanthoids) have undergone extensive parallel and convergent evolution, and they have frustrated attempts to produce a natural generic classification based on macromorphologic characteristics alone. Although some workers have aggregated species into a few large genera (e.g., J. T. Mickel 1979b), most tend to recognize smaller segregate genera based on a combination of morphologic, chromosomal, and biochemical data. The latter approach seems to provide a more useful, evolutionarily informative classification and is the one adopted here. Aspidotis and Notholaena are maintained here as distinct from Cheilanthes , and three recently described genera ( Argyrochosma , Astrolepis , and Pentagramma ) have been incorporated into the treatment. The reasons for these changes in generic circumscription are discussed under the individual genera.

Species within checklist: Flora of the National Park Service, Intermountain Region || << 1 - 50 taxa >>
Adiantum capillus-veneris
Image of Adiantum capillus-veneris
Adiantum pedatum
Image of Adiantum pedatum
Argyrochosma fendleri
Image of Argyrochosma fendleri
Argyrochosma jonesii
Image of Argyrochosma jonesii
Argyrochosma limitanea
Image of Argyrochosma limitanea
Argyrochosma microphylla
Image of Argyrochosma microphylla
Astrolepis cochisensis
Image of Astrolepis cochisensis
Astrolepis integerrima
Image of Astrolepis integerrima
Astrolepis sinuata
Image of Astrolepis sinuata
Astrolepis windhamii
Image of Astrolepis windhamii
Bommeria hispida
Image of Bommeria hispida
Cheilanthes alabamensis
Image of Cheilanthes alabamensis
Cheilanthes bonariensis
Image of Cheilanthes bonariensis
Cheilanthes covillei
Image of Cheilanthes covillei
Cheilanthes eatonii
Image of Cheilanthes eatonii
Cheilanthes feei
Image of Cheilanthes feei
Cheilanthes fendleri
Image of Cheilanthes fendleri
Cheilanthes horridula
Image of Cheilanthes horridula
Cheilanthes kaulfussii
Image of Cheilanthes kaulfussii
Cheilanthes lendigera
Image of Cheilanthes lendigera
Cheilanthes lindheimeri
Image of Cheilanthes lindheimeri
Cheilanthes parryi
Image of Cheilanthes parryi
Cheilanthes pringlei
Image of Cheilanthes pringlei
Cheilanthes tomentosa
Image of Cheilanthes tomentosa
Cheilanthes villosa
Image of Cheilanthes villosa
Cheilanthes wootonii
Image of Cheilanthes wootonii
Cheilanthes wrightii
Image of Cheilanthes wrightii
Cheilanthes yavapensis
Image of Cheilanthes yavapensis
Myriopteris aurea
Image of Myriopteris aurea
Myriopteris covillei
Image of Myriopteris covillei
Myriopteris fendleri
Image of Myriopteris fendleri
Myriopteris gracilis
Image of Myriopteris gracilis
Myriopteris lendigera
Image of Myriopteris lendigera
Myriopteris lindheimeri
Image of Myriopteris lindheimeri
Myriopteris parryi
Image of Myriopteris parryi
Myriopteris rufa
Image of Myriopteris rufa
Myriopteris tomentosa
Image of Myriopteris tomentosa
Myriopteris windhamii
Image of Myriopteris windhamii
Myriopteris wootonii
Image of Myriopteris wootonii
Myriopteris wrightii
Image of Myriopteris wrightii
Myriopteris yavapensis
Image of Myriopteris yavapensis
Notholaena aliena
Image of Notholaena aliena
Notholaena grayi
Image of Notholaena grayi
Notholaena greggii
Image of Notholaena greggii
Notholaena lemmonii
Image of Notholaena lemmonii
Notholaena nealleyi
Image of Notholaena nealleyi
Notholaena neglecta
Image of Notholaena neglecta
Notholaena standleyi
Image of Notholaena standleyi
Pellaea atropurpurea
Image of Pellaea atropurpurea
Pellaea breweri
Image of Pellaea breweri
The National Science Foundation
Development supported by National Science Foundation Grants (DBI 9983132, BRC 0237418, DBI 0743827, DBI 0847966)
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