Perennials; not scapose; pubescent or glabrous, trichomes simple, with stalked, forked or dendritic ones. Stems erect [ascending], often branched distally. Leaves basal and cauline; petiolate or sessile; basal (withered with age), rosulate, petiolate, blade margins entire, dentate to sinuate, or runcinate; cauline subsessile or sessile [shortly petiolate], blade margins entire [dentate to sinuate]. Racemes (several-flowered, sometimes secund, lax), considerably elongated in fruit. Fruiting pedicels erect, descending, ascending, or divaricate-ascending, slender, (glabrous). Flowers (usually actinomorphic, rarely zygomorphic, cup-shaped); sepals erect, oblong to ovate, lateral pair slightly to strongly saccate basally, (usually pubescent, rarely glabrous); petals white (lavender or purple apically), spatulate to oblanceolate, (± longer than sepals), claw undifferentiated from blade, (apex obtuse); stamens slightly tetradynamous; filaments dilated basally; anthers ovate to oblong, (apex obtuse); nectar glands confluent, subtending bases of stamens. Fruits siliques, sessile or shortly stipitate, linear, not torulose, straight or curved, terete or latiseptate; valves each with prominent or obscure midvein, glabrous or pubescent; replum rounded; septum complete; ovules 40-250 per ovary; style distinct or obsolete; stigma capitate. Seeds uniseriate or biseriate, plump or flattened, winged or not, oblong to ovate; seed coat (reticulate-areolate), mucilaginous or not when wetted; cotyledons accumbent or incumbent. x = 8.