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Grammitidaceae
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Alan R. Smith in Flora of North America (vol. 2)
Plants perennial, mostly small, on rock or commonly epiphytic [rarely terrestrial]. Stems long- to short-creeping or suberect, usually unbranched, bearing scales [rarely scales absent], solenostelic (having phloem on both sides of xylem) to dictyostelic (having complex nets of xylem). Leaves erect, arching, or pendent, monomorphic [rarely with specialized fertile areas], less than 50 cm [rarely longer], usually scaleless throughout. Petioles often dark-colored and wiry, commonly terete, usually less than 2 mm diam., articulate or not articulate, with 1 or 2 vascular strands. Blades simple and entire to commonly pinnatifid or 1-pinnate, rarely 2-pinnate or more divided, glabrous or commonly bearing hairs, especially on petioles and rachises; hairs tan to dark reddish brown [or transparent], unicellular to multicellular; rachises often dark-colored, not grooved adaxially. Veins free [to anastomosing in simple patterns]; hydathodes present or absent, sometimes obscured by lime dots adaxially. Sori abaxial on veins, round to oblong [occasionally elongate]; paraphyses present or absent, these glandular or hairlike; sporangia with stalk of 1 row of cells; indusia absent. Spores greenish, tetrahedral-globose, trilete, surface commonly papillate. Gametophytes greenish, borne aboveground, ribbon-shaped, sometimes bearing multicellular gemmae. Circumscription of genera is unsettled and in some cases arbitrarily based on blade dissection. Generic limits need redefinition and are the subject of current study. The family description above encompasses the worldwide variation. Relationships of the family have generally been thought to be with the Polypodiaceae with which the family is sometimes combined, but that is open to question. Most species are epiphytic, and many occur in cloud forests at middle and high elevations.

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Ceradenia albidula
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Ceradenia argyrata
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Ceradenia aulaeifolia
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Ceradenia brunneoviridis
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Ceradenia capillaris
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Ceradenia clavipila
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Ceradenia curvata
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Ceradenia discolor
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Ceradenia farinosa
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Ceradenia fendleri
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Ceradenia fucoides
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Ceradenia intonsa
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Ceradenia intricata
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Ceradenia jungermannioides
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Ceradenia kalbreyeri
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Ceradenia knightii
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Ceradenia kookenamae
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Ceradenia margaritata
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Ceradenia mayoris
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Ceradenia melanopus
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Ceradenia meridensis
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Ceradenia nubigena
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Ceradenia oidiophora
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Ceradenia pearcei
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Ceradenia pilipalaea
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Ceradenia pilipes
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Ceradenia podocarpa
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Ceradenia pruinosa
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Ceradenia sacksii
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Ceradenia semiadnata
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Ceradenia similis
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Ceradenia spixiana
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Ceradenia tristis
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Ctenopteris blechnoides
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Ctenopteris flabelliformis
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Ctenopteris seemannii
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Lellingeria apiculata
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Lellingeria depressa
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Lellingeria dissimulans
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Lellingeria flagellipinnata
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Lellingeria hartii
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Lellingeria humilis
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Lellingeria isidrensis
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Lellingeria labiakii
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Lellingeria longeattenuata
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Lellingeria major
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Lellingeria micula
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Lellingeria militaris
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Lellingeria myosuroides
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Lellingeria pendula
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The National Science Foundation
Development supported by National Science Foundation Grants (DBI 9983132, BRC 0237418, DBI 0743827, DBI 0847966)
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