Subshrubs or shrubs, 10-80+[-150] cm. Stems usually 1, procumbent to erect, usually branched, glabrous [hairy]. Leaves mostly cauline; alternate; petiolate or sessile; blades ± obovate [oblong to lanceolate or linear] (bases sometimes ± clasping), (1-)2-3-pinnately lobed (lobes cuneate to linear), ultimate margins dentate [entire], faces glabrous [hairy]. Heads radiate [discoid], borne singly or in open, corymbiform arrays. Involucres hemispheric or broader, [6-]10-18[-22+] mm diam. Phyllaries persistent, 28-45+ in 3-4 series, distinct, oblanceolate or ovate to lance-deltate or lanceolate (not keeled abaxially), unequal, margins and apices (stramineous to brown) scarious (tips of inner often ± dilated). Receptacles convex to conic, epaleate. Ray florets 12-35+, pistillate, fertile; corollas usually white, sometimes yellow or pink, laminae ± ovate to linear. Disc florets [50-]80-150+, bisexual, fertile; corollas yellow [red, purple], tubes ± cylindric (not basally dilated, ± gland-dotted), throats campanulate, lobes 5, deltate (without resin sacs). Cypselae dimorphic: outer (ray) 3-angled, each angle usually ± winged (wings not spine-tipped); inner (disc) compressed-prismatic (± quadrate, sometimes 2 angles winged, wings not spine-tipped); all ± ribbed or nerved, faces usually glabrous, sometimes gland-dotted between ribs (pericarps without myxogenic cells or resin sacs; embryo sac development bisporic); pappi 0 (cypselar wall tissue sometimes produced as teeth, crowns, or oblique tubes similar in texture to cypselar wings). x = 9.