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Phyla cuneifolia (Torr.) Greene  
Family: Verbenaceae
Wedgeleaf, more...fogfruit, wedgeleaf fogfruit
[Lippia cuneifolia (Torr.) Steud., moreLippia cuneifolia var. incisa Blank., Zappania cuneifolia]
Phyla cuneifolia image
Frankie Coburn
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The Morton Arboretum
Perennial subshrub or herb 5 - 10 cm tall Stem: trailing but not rooting at nodes, densely hairy. Leaves: opposite, 1.5 - 4 cm long, 3 - 8 mm wide, narrow and inversely egg-shaped with a wedge-shaped base and a broadly rounded tip, with one to three teeth per side. Flowers: borne in dense axillary spikes, subtended by abruptly pointed bracts, with a small and membranous calyx and stamens shorter than to slightly longer than the corolla tube. The slender corolla has a tube slightly longer than the calyx and four unqual spreading lobes, appearing somewhat two-lipped. Fruit: two nutlets surrounded by or joined to the calyx.

Similar species: Phyla species have axillary inflorescences, four-lobed corollas and fruit separating into two nutlets. Phyla lanceolata differs by having lance- to egg-shaped leaves with pointed leaf tips and five to eleven teeth per side.

Flowering: May to September

Habitat and ecology: Introduced from the western United States, this species is only found in Porter County. Elsewhere, it grows in disturbed areas.

Occurence in the Chicago region: non-native

Etymology: Phyla means tribe. Cuneifolia means "having wedge-shaped leaves."

Author: The Morton Arboretum

Kearney and Peebles 1969, McDougall 1973
Duration: Perennial Nativity: Native Lifeform: Subshrub General: Herbaceous perennials from a woody base; stems trailing along the ground and rooting at the nodes, 20-100 cm long, puberulent, square in cross section. Leaves: Opposite and sessile, clustered on short branches at stem nodes; blades oblanceolate to obovate, 1-3 cm long, thickish, with a conspicuous midrib, margins with 1-4 pairs of teeth close to the leaf tip, surfaces canescent with long, soft hairs. Flowers: White-lavender, clustered in dense capitate spikes, these globose in flower and elongating to ovoid in fruit, on peduncles from the leaf axils; peduncles shorter than or up to 2 times as long as subtending leaves; each flower subtended by a broad bractlet, these imbricate where visible at the bottom of the spike; calyxes campanulate, 2-lipped, 4-toothed, and strigose; corollas white to pink to purple, 4 mm long, 2-lipped, with 4 lobes. Fruits: Schizocarps oval, separating into 2 nutlets. Ecology: Found in gravelly soils, near stream beds and playas, from 4,000-7,000 ft (1219-2134 m); flowers June-August. Distribution: SD to WY and UT, south to TX, NM, AZ, and MEX. Notes: This distinctive trailing herb is uncommon, though present in most Arizona and New Mexico counties. The stems trail along the ground, and each stem node has a cluster of long, narrow leaves with prominent midveins and a few teeth near the slightly widened tips. Some of the stem nodes also have a round cluster of light purple flowers elevated above the leaves on a slender peduncle. These distinctive capitate heads with the little bilabiate flowers could be confused with Valeriana. However, Phyla has corollas with 4 lobes, while Valeriana has corollas with 5 lobes and a distinctive smell. Ethnobotany: Poultice of plants applied to spider bites. Synonyms: Zapania cuneifolia, Lippia cuneifolia, Lippia cuneifolia var. incisa Editor: LCrumbacher2012, AHazelton 2015 Etymology: Phyla comes from the Greek phyle, "tribe," probably from the flowers being tightly clustered in heads, and cuneifolia means with leaves tapered to the base.
Vascular plants of NE US and adjacent Canada
A western sp., is intr. at St. Louis and may be expected in our region. It differs from both of the following spp. in its narrow, linear-oblanceolate or narrowly cuneate lvs 4-10 times as long as wide, less than 1 cm wide, with 1-3 teeth on each side toward the summit.

Gleason, Henry A. & Cronquist, Arthur J. 1991. Manual of vascular plants of northeastern United States and adjacent Canada. lxxv + 910 pp.

©The New York Botanical Garden. All rights reserved. Used by permission.
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Frankie Coburn
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Greg Goodwin
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Kirstin Phillips
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Kirstin Phillips
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Development supported by National Science Foundation Grants (DBI 9983132, BRC 0237418, DBI 0743827, DBI 0847966)
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