Leaves sometimes appearing whorled at ends of branches. Flowers erect to drooping; peduncle smooth to rugulose; perianth white, yellow, yellow-green, pink, or red to maroon, often glandular; outer tepals often bractlike and reduced, sometimes ciliolate, inner tepals often much larger and ligulate, sometimes only slightly larger and broadly obtuse, innermost often reduced, sometimes transitional to stamens; pistils orbicular to deltoid, styles narrow and acute, often recurved. Fruits dark brown, woody to leathery at maturity; peduncle sometimes thickened. Seeds glossy, dark brown to tawny or golden. x = 13, 14.