Dryopteris spp.
Family: Dryopteridaceae
Project: Southwest Biodiversity Consortium
Plants terrestrial, rarely on rock. Stems short-creeping to erect, stolons absent. Leaves monomorphic, green through winter or dying back in winter. Petiole ca. 1/4--2/3 blade length, bases swollen or not; vascular bundles more than 3, arranged in an arc, ± round in cross section. Blade deltate-ovate to lanceolate, 1--3-pinnate-pinnatifid, gradually reduced distally to pinnatifid apex, herbaceous to somewhat leathery. Pinnae not articulate to rachis, segment margins entire, crenate, or serrate, spinulose or not; proximal pinnae reduced (several pairs), same size as or enlarged relative to more distal pinnae, sessile to petiolulate, equilateral or often inequilateral with pinnules on basiscopic side longer than those on acroscopic side; costae adaxially grooved, grooves continuous from rachis to costae to costules; indument of linear to ovate scales abaxially, also sometimes with glands, blades ± glabrous adaxially. Veins free, forked. Sori in 1 row between margin and midrib, round; indusia round-reniform, attached at narrow sinus, persistent or caducous. Spores brownish, coarsely rugose or with folded wings. x = 41. The relationships of the North American species are reasonably well understood, but species identifications are complicated by the frequent presence of hybrids in field populations. Sterile hybrids can be distinguished from fertile species by their misshapen spores and intermediate morphology. They are not included in the key, but they may be identified as to parentage by combinations of characters in the key (e.g., marginal sori for Dryopteris marginalis , narrow blades for D . cristata ). Relationships are shown in the accompanying reticulogram.

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