Arctium spp.
Family: Asteraceae
Project: Southwest Biodiversity Consortium
Biennials or (monocarpic) perennials, 50-300 cm; herbage not spiny. Stems erect, openly branched, branches ascending. Leaves basal and cauline; long-petiolate; gradually smaller distally; blade margins entire or dentate (pinnately lobed or dissected), faces abaxially resin-gland-dotted, adaxially often tomentose. Heads discoid, in leafy-bracted racemiform to paniculiform or corymbiform arrays. ( Peduncles 0 or 1-9 cm.) Involucres spheric to ovoid. Phyllaries many in 9-17 series, outer and mid narrowly linear. bases appressed, margins entire. apices stiffly radiating, hooked-spiny tipped, inner linear, ascending or erect, straight tipped. Receptacles ± flat, epaleate, bearing subulate scales. Florets (5-)20-40+; corollas pink to ± purple, glabrous or glandular-puberulent, tubes elongate. throats campanulate. lobes narrowly triangular, ± equal; anther bases tailed, apical appendages ovate, obtuse to acute; style branches: fused portions distally hairy-ringed, distinct portions oblong, acute or obtuse . Cypselae obovoid. ± compressed, rough or ribbed, glabrous, attachment scars basal; pappi falling, of many bristles in 2-4 series . x = 18. At maturity the dry heads of Arctium species are readily caducous with the enclosed cypselae, and the hooked phyllary tips cling easily to fur or fabrics. Animal dispersal is a major factor in the spread of burdock species across North America. The burs are a major problem when they become entangled in the wool of sheep and fur of dogs and other animals. Published chromosome reports for Arctium other than n = 18 are probably in error because of difficulty in interpretation of somatic chromosomes (R. J. Moore and C. Frankton 1974).

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