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Family:
Asteraceae
rough cocklebur, more..., Canada cockleburr, cocklebur, cockleburr, common cocklebur, rough cockleburr, (Spanish: cadillo)
[Xanthium acerosum Greene, more, Xanthium californicum Greene, Xanthium californicum var. rotundifolium Widder, Xanthium campestre Greene, Xanthium canadense P. Mill., Xanthium cavanillesii Schouw, Xanthium cenchroides Millsp. & Sherff, Xanthium commune Britt., Xanthium echinatum Murr., Xanthium glanduliferum Greene, Xanthium italicum Moretti, Xanthium macounii Britt., Xanthium oligacanthum Piper, Xanthium oviforme Wallr., Xanthium pensylvanicum Wallr., Xanthium saccharatum Wallr., Xanthium speciosum Kearney, Xanthium strumarium ssp. italicum (Moretti) D. Löve, Xanthium strumarium var. canadense (P. Mill.) Torr. & A. Gray, Xanthium strumarium var. oviforme (Wallr.) M.E. Peck, Xanthium strumarium var. pensylvanicum (Wallr.) M.E. Peck, Xanthium varians Greene]
 Max Licher |
Plants 10-80(-200) cm; nodal spines 0. Leaves: petioles 20-100(-140+) mm; blades suborbiculate to ± pentagonal or deltate, 4-12(-18+) × 3-10(-18+) cm, sometimes palmately 3-5-lobed, abaxial faces green, hirtellous. Burs 10-30+ mm. 2n = 36. Flowering Jul-Oct. Damp or seasonally wet, often alkaline, soils, waste places, margins of agriculture; 10-2000 m; Alta., B.C., Man., N.B., N.S., Ont., P.E.I., Que., Sask.; Ala., Alaska, Ariz., Ark., Calif., Colo., Conn., Del., D.C., Fla., Ga., Idaho, Ill., Ind., Iowa, Kans., Ky., La., Maine, Md., Mass., Mich., Minn., Miss., Mo., Mont., Nebr., Nev., N.H., N.J., N.Mex., N.Y., N.C., N.Dak., Ohio, Okla., Oreg., Pa., R.I., S.C., S.Dak., Tenn., Tex., Utah, Vt., Va., Wash., W.Va., Wis., Wyo.; Mexico; Central America; South America; widely introduced in Old World. Recognition of a dozen or more taxa (treated as species, subspecies, varieties, and/or forms) has been proposed for plants treated together here as Xanthium strumarium. Bases for the various taxa mostly involved subtle differences in the burs.
Plant: Monoecious annual forb to 80 cm; herbage glandular and scabrous Leaves: leaves alternate, long-petiolate, blades 5-15 cm wide and long, often 3-lobed, margins serrate INFLORESCENCE: primary inflorescence a head, each resembling a flower; staminate heads in clusters, involucre 0, receptacle chaffy; pistillate heads clustered below staminate heads, 2-flowered, 2-beaked, spiny, bur-like; phyllaries 0 or minute; chaff scales many, spirally arrayed, fused below, free tips spiny, hooked Flowers: Staminate flowers many; corolla translucent; filaments fused, attached to corolla tube base, anthers free; ovary slender; Pistillate flowers generally 2 per head; corolla 0; style branches long Fruit: Fruit: 2, enclosed in bur, germinating in successive years; bur cylindric to barrel-shaped; spines generally stout, ± glandular; pappus 0 Misc: Disturbed areas; generally < 500 m.; Jul-Oct Notes: Flower has no corolla or pappus.Stamens monodelphous.Style unbranched.The spines on the fruit may be glandular. References: McDougall; Seed plants of Northern Arizona. Hickman, ed.; The Jepson Manual. ASU specimans
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